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25Diplommatina polypleurisBenson, 1857 Fig. 19 A, B Diplommatina polypleuris Benson, 1857: 203. Type locality: ad Nancla. Theobald 1858: 318; Pfeiffer, 1858: 11. Godwin-Austen 1868: 83, pl. 3, fig. 1. Theobald and Stoliczka 1872: 330. Hanley and Theobald 1875: 56, pl. 140, fig. 10. Pfeiffer 1875: 73. Theobald 1876: 42. Nevill 1878: 285. Godwin-Austen 1884: pl. 45, figs 1, 1 a. Godwin-Austen 1886: 176. Gude 1921: 328. Diplommatina polypleuris var. Godwin-Austen, 1870: 4, pl. 1, figs 4, 4 a. Type locality: North Cachar and north Jaintia hills [mountainous region in Assam and Meghalaya state, India]. Diplommatina polypleuris var. minuta Godwin-Austen, 1876: 178 (nomen nudum). Diplommatina polypleuris var. minuta Godwin-Austen, 1886: 176 [in the name usage list]. Diplommatina (Eudiplommatina) polypleuris — Kobelt and Möllendorff 1898: 137. Diplommatina (Diplommatina) polypleuris — Kobelt 1902: 441. Preece et al. 2022: 86, fig. 34 e. Type material examined. • Syntype. UMZC I.102640 (1 shell), no locality data; R. McAndrew coll. Other material. • NHMUK 1903.7.1.2314 (2 shells) from Jawai, Jaintia; Godwin-Austen coll. NHMUK 1903.7.1.2177 (1 shell) from Khasi (figured in Godwin-Austen 1884: pl. 45, figs 1, 1 a); Godwin-Austen coll. NHMUK 1903.7.1.2269 (24 shells) from Shillong, Khasi Hills; Godwin-Austen coll. NHMUK 1905.5.5.49 (12 shells) from North Scarp, Khasi Hills; W. T. Blanford coll. UMZC I.102645 (1 shell), no locality data; R. McAndrew coll. NHMUK 1891.3.17.728 –730 (3 shells; Fig. 19 B) labelled as D. polypleuris var. from Nattoung; Sowerby coll. NHMUK 1903.7.1.2252 (5 shells; Fig. 19 A) labelled as D. polypleuris var. minuta from Nongtung, Khasi; Godwin-Austen coll. NHMUK 1903.7.1.2262 (1 shell) labelled as D. polypleuris var. minuta from Japvo Peak, Naga; Godwin-Austen coll. NHMUK 1903.7.1.263 (1 shell) labelled as D. polypleuris var. minuta from Munken Valley, Jaintia; Godwin-Austen coll. Diagnosis. Shell dextral, ovate-fusiform, translucent, whitish to yellowish. Whorls ~ 5, rounded; spire depressed conical; suture wide, deep. Teleoconch with strong, densely and equally spaced radial ribs throughout. Last three whorls nearly equal in width; penultimate whorl slightly wider than others. Aperture rounded; columellaris weak, indistinct. Peristome expanded, double, multi-layered; outer peristome expanded beyond inner peristome. Umbilicus narrowly open. Distribution. This species was originally known from its type locality in Nanclai Poonjee, Assam, India, and was later recorded from the Dafla Hills, North Khasi Hills, and Darjeeling (Hanley and Theobald 1870; Gude 1921; Preece et al. 2022). In Myanmar, it has been reported from Nattoung, Sandoway, which likely corresponds to the Thandwe District area in Rakhine State (Theobald and Stoliczka 1872; Preece et al. 2022). Remarks. Godwin-Austen (1870: 4) recognised specimens from ‘ North Cachar and North Jaintia Hills’ as a distinct variety, but did not provide a name for it. Subsequently, he introduced the name var. minuta for specimens from ‘ Shengorh and Torúpútú Peaks’ (Godwin-Austen 1876: 178). However, this nomination lacks a definition, indication, description, and any reference to earlier work; therefore, this varietal name was not made available (ICZN 1999: Art. 12). Later, in the list of name usages under D. polypleuris, Godwin-Austen (1886: 176) stated ‘ from N. Jaintia is distinct; I have named it minuta ’, with an explicit indication referring to the published description in Godwin-Austen (1870). It seems that the name var. minuta was made available from Godwin-Austen (1886). Regarding the type series, the NHM holds one lot, NHMUK 1903.7.1.2252 (4 shells), registered as syntypes of D. polypleuris minuta. However, the recorded locality is from ‘ Nongtung, Khasi’, which does not match the locality ‘ North Jaintia Hills’ associated with the original description. Therefore, this specimen lot is unlikely to be the syntype, since it neither originates from the type locality nor appears to be part of the type series examined by Godwin-Austen. The specimen figured in Preece et al. (2022: fig. 34 e), labelled as D. polypleuris, appears to have prominent radial ribs and a distinct horizontal palatalis, whereas the specimens from Nattoung (Fig. 19 B), referred to as D. polypleuris var., tend to have weaker radial ribs and a conspicuous vertical plica. These differences suggest that the two are unlikely to represent the same taxon.
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ประเด็นที่เกี่ยวข้อง: Diatoms and Algae Research · Aquatic Invertebrate Ecology and Behavior · Aquatic Ecosystems and Phytoplankton Dynamics
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Piyoros Tongkerd · Arthit Pholyotha · Supunya Annate · Chirasak Sutcharit · Chulalongkorn University
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